Jonathan Balcombe is a biologist, author and life-long animal advocate. He has a Ph.D. in ethology from the University of Tennessee, where he studied communication in bats. He has published over 50 journal articles and book chapters ranging from turtle nesting behavior to the ethics of animal dissection. His 2006 book “Pleasurable Kingdom” is the first in-depth examination of animals’ capacity to enjoy life. His subsequent books “Second Nature” and “The Exultant Ark” also present animals in a new light and presage a revolution in the human-animal relationship. His latest book, the New York Times bestseller “What a Fish Knows,” explores the private lives of the planet’s most misunderstood and maligned vertebrates. Balcombe is Director for Animal Sentience with The Humane Society Institute for Science and Policy, based in Washington, D.C. He is also a founding editor of Animal Sentience, the first scholarly journal of animal feeling.